Leaves and Stem Leaf features to look out for Leaves are generally speaking broad, flat, and thin.
Many of the bolded characters in the characterization above are apomorphies of more or less inclusive clades of streptophytes along the lineage leading to the embryophytes, not apomorphies of crown-group embryophytes per se. All groups below are crown groups, nearly all are extant.
Characters mentioned are those of the immediate common ancestor of the group,  contains explanatory material, features common in clade, exact status unclear. B class], mitochondrial rps2 gene lost.
G , superposed; loss of introns in RPB2 gene d copy [? For the complex patterns of variation in a number of characters in this part of the tree, see the Gentianales page.
For the relationships of Lamiales, see discussion under Gentianales. In all node characterizations, boldface denotes a possible apomorphy, Note that the particular node to which many characters, particularly the more cryptic ones, should be assigned is unclear.
This is partly because homoplasy is very common, in addition, basic information for all too many characters is very incomplete, frequently coming from taxa well embedded in the clade of interest and so making the position of any putative apomorphy uncertain.
Then there are the not-so-trivial issues of how character states are delimited and ancestral states are reconstructed see above. Estimates of the age of crown-group Lamiales are about Table S277 m.
Lamiales contain ca Most of this diversity is concentrated in families whose members are herbaceous to shrubby and have rather large, monosymmetric flowers, and about half have fruits with many rather small seeds Simsand although about half the species have only eight or fewer seeds per fruit, they are not very big.
For a useful general discussion, including suggestions of apomorphies for some clades, see Soltis et al. Endress a suggested that a key innovation somewhere in Lamiales was tenuinucellate ovules.
Ovule number is notably variable in the basal clades and will be difficult to optimize. The four clades that are successively sister to other Lamiales either lack iridoids or have iridoids distinctively different Oleaceae from those in the other members of the clade, so iridoid re aquisition is pegged well within Lamiales; whether or not Carlemanniaceae have iridoids is unknown.
Monosymmetry is unlikely to be plesiomorphic in the order c. Floral evolution in basal Lamiales is not simple, and where changes in floral meristicity and floral symmetry are to be placed on the tree is unclear. Endress b suggested that families such as Orobanchaceae, Lamiaceae and Acanthaceae form a clade with strongly monosymmetric flowers that mostly lack a staminode, but such a grouping is not obviously consistent with the relationships being recovered.
Confirmation of the phylogenetic positions of Carlemanniaceae, placed sister to Oleaceae, and of Plocospermataceae, as well as studies of their anatomy, chemistry, floral development, etc. Thus, given their position, one might expect Carlemanniaceae to lack iridoids - at least, to lack route II decarboxylated iridoids - and to have only a single micropylar endosperm haustorium.
As might be anticipated, there is little morphological support for internal nodes in much of Lamiales and also for several of the families, and this is likely to be true whatever the relationships in the order. For the complex pattern of variation in a number of other characters in this part of the tree, see the Gentianales page.
Carnivory, direct or indirect, has arisen three times in Lamiales Lentibulariaceae, Byblidaceae, Plantaginaceaeand Martyniaceae may also be carnivorous. For other characters that may clarify the relationships of Lamiales, see Gentianales.
A great deal of work on characterising iridoids and understanding their distribution in Lamiales has been carried out by S.
The presence of cornosides and iridoids in Lamiales is largely mutually exclusive, except in Martynia louisiana Jensena, b.3a pharmaceuticals: 3g life sciences rutadeltambor.com: 3m espe dental professionals: 3m india ltd formerly birla: 4 care lifescience pvt ltd: a d pharma: a one pharma rutadeltambor.com Team Leader Executive Summary - Executive Summary Team Leader offers top notch training to professional engineering fields, providing the skills and information needed by employees to maintain high standards within our corporation.
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